The majority of records pertain to the rio Guaporé (Río Iténez in Bolivia), the main affluent of the rio Mamoré, which drains the vast tropical wetland known as the Pantanal, and the upper rio Paraguai, which also has its headwaters in the Pantanal but flows in the opposite direction. The Guaporé and Paraguai are connected due to rising water levels during the annual wet season.