During collections of the type series stream width was found to vary seasonally but was never wider than 9.1 metres. Water depth was 10-40 cm, flow rate 20-70 cm/s−1, and substrates comprised small to medium-sized rocks.

Water temperature was seasonally variable but always within the range 20-26°C/68-78.8°F, pH was 6.5-7.9, oxygen 7.4-8.5 mg/l−1, ammonia, nitrate and total iron were 0.01, < 0.03 and < 0.5 mg/l−1, respectively, and alkalinity was aro...