Benito

Described from northern Gabon, close to 'Zoumoukou', which is seeen on modern maps as 'Zomoko', but has a much wider distribution extending into Equatorial Guinea (upper parts of some Benito River tributaries), and southern Cameroon (Ntem River), as well as the Ivindo and Okano drainages in Gabon, the latter pair both tributaries of the much larger Ogooué system.

Described from Río Muni, Equatorial Guinea and appears endemic to south-western parts of the country, with additional collections having occurred in the Río Aye, Río Ecucu and lower Río Benito drainages.