Garra species are usually included in the subfamily Labeoninae/Cyprininae (name varies with author) which by current thinking is further divided into several tribes and subtribes (Stiassny and Getahun, 2007). The putatively monophyletic subtribe Garraina comprises Garra plus numerous other genera including the well-known Crossocheilus and Epalzeorhynchos. Within this division their closest relatives are members of Discogobio, Discocheilus and Placocheilus (the lat...

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