Henicorhynchus entmema

Recorded from the middle to lower Mekong River basin in Laos, Thailand, and Cambodia, plus the Chao Phraya and Mae Klong watersheds in central and western Thailand.

Type locality is 'Thailand: Chiang Rai Province: Mekok River near Chiang Rai'.

This species is considered to be one of the most abundant species in the massive seasonal migrations of cyprinid fishes that occur in the mainstream Mekong river in December to February and May to July. Otherwise, it has been recorded in rivers and streams of all sizes down to around 2-3 m in width, with slow to rapid flow, and is known to enter inundated forests during the wet season.

H. entmema can be distinguished from congeners by the following combination of characters: possession of 32-33 lateral line scales; 5½ scale rows between lateral line and base of dorsal-fin; body relatively slender, depth fits 3.7-4.0 times in SL; snout often projecting strongly; body plain silver in colour with no dark marking on caudal peduncle.

Species currently included in Gymnostomus have previously been placed in a variety of genera including Henicorhynchus, Cirrhinus, Bangana, and Labeo in the last few decades alone. However, Gymnostomus is a senior subjective synonym of Henicorhynchus, thus rendering the latter invalid, while Gymnostomus is easily distinguished from Cirrhinus by possession of 8-9 (vs. 10-15) branched dorsal-fin rays.

Some confusion remains, however, since a recent phylogenetic study (Yang et al., 2012) considered Henicorhynchus to be valid, and members were not recovered in the same evolutionary lineage as Indian Gymnostomus species, suggesting that they are not closely related. It is unclear whether further taxonomic changes will be required in the future in order to reflect this disparity, but here on SF we currently follow Kottelat (2013) since it was published more recently and has been widely accepted, thus Henicorhynchus is a synonym of Gymnostomus.

Gymnostomus species are considered members of the tribe Labeonini within the putative cyprinid subfamily Cyprininae or simply the subfamily Labeoninae (name varies with author). According to the most recent phylogenetic research, this grouping is further divided into four subtribes; Labeoina, Garraina, Osteochilina, and Semilabeoina (Yang et al., 2012). Among these, Gymnostomus is included in the Labeoina alongside Labeo, Bangana sensu stricto (which includes the genus Nukta), Cirrhinus sensu stricto, Cirrhinus microlepis (which is of a different genetic lineage to other Cirrhinus species), and Incisilabeo. As noted above, however, this refers only to the Indian species of Gymnostomus, with the Southeast Asian species placed in the Osteochilina assemblage.

Some Gymnostomus species are common in mainstream rivers at certain times of year and extremely important in local fisheries during these periods.