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Heterandria formosa

Least Killifish

(Girard, 1859)

Last revised 2026Edit this page

On this page11 sections
Heterandria formosa male
Heterandria formosa maleHippocampus-Bildarchiv

Quick factsEdit

Max size
20mm SL0.8in SL
Aquarium size
not recorded
Temp
20–26°C68–79°F
pH
7.0–8.0
Hardness
5–20dGH

Difficulty6 of 8 measures

Easy

How the rating is worked out

Space
small tank
Water
needs a steady pH
Temp
flexible
Diet
unknown
Temperament
peaceful
Social
needs to be in a group
Compatibility
choose tankmates carefully

Other demands

Each of these can raise the rating on its own, whatever the measures above say.

Hardiness
unknown

The least killifish, Heterandria formosa, is a tiny livebearer from the United States, found in heavily vegetated margins of still or sluggish water. The smallest known livebearing fish, it is peaceful but not a community fish in the general sense, as the males reach only 2 cm and many commonly kept species will eat them. It is sometimes confused with the much more aggressive mosquito fish of the genus Gambusia.

Distribution

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Type locality is 'Charleston, South Carolina and Palatka, Florida, U.S.A.'

It's usually found in shallow, heavily-vegetated marginal areas of still or sluggish bodies of water. It can also be found in brackish conditions in some areas.

Aquarium size

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Standard length
20mm SL0.8in SL
Aquarium base
not recorded

A pair of these can be kept in a tank as small as 12" x 8" x 8" (30cm x 20cm x 20cm) - 12.5 litres.

Maintenance

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A gently-filtered, heavily planted setup suits it best. Other decor can be added as you wish. Water flow must be kept to a minimum.

Water conditions

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Temperature
20–26°C68–79°F
pH
7.0–8.0
Hardness
5–20dGH

Temperature: It's a hardy species, being subject to a large range of temperatures in its natural waters. Somewhere between 68-78°F (20-26°C) is best in an aquarium.

pH: 7.0-8.0

Hardness: 5-20°H

Unfussy and omnivorous, it will accept most foods offered. It's particularly fond of small live or frozen varieties such as brine shrimp or Daphnia, and the diet should contain a good proportion of these. It will also browse on algae, so try to ensure it receives some vegetable matter in the diet. In the absence of algae crushed spirulina flakes work well.

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Behaviour and compatibility

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Not a community fish in the general sense, although it's very peaceful. Males in particular are so tiny that they will be seen as a snack by many of the commonly kept species. It will also not compete well with more vigorous tankmates. Other similarly-sized, non-predatory species such as pencil fish, dwarf Corydoras, Otocinclus and freshwater shrimp are the best choices.

While males can show a little aggression when competing for females, physical damage is very rare. It's best to maintain it in a small colony.

Sexual dimorphism

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Males are much smaller than females and possess an enormous gonopodium.

Reproduction

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Very easy. It's nigh on impossible to prevent it breeding if both sexes are present in an aquarium. Water parameters are unimportant, provided they are within the ranges stated above. The tank should be planted heavily. It's best kept in a small group with several of each sex present for bredding purposes.

The gestation period is around 4 weeks. The species has a slightly different method of fry production to most other livebearers, involving a process known as 'superfoetation'. This is defined as 'formation or development of a second foetus when one is already present in the uterus'. Fry at different stages of development can therefore be present in the uterus of the fish at any given time. In addition, the egg yolks of the species are nutritionally poor, and the developing fry derive much of their nourishment via organs that function in a similar way to the placenta of mammals. As a result of this process, fry are dropped continually rather than in defined broods. You'll see a few fry appearing every day or two if you have more than one female in the tank. They are quite large at birth and can accept powdered dry foods and Artemia nauplii straight away. The adults will not usually harm them.

Notes

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This species is also referred to as 'mosquito fish', 'dwarf livebearer' or 'dwarf top minnow' in aquarium literature.

, H. formosa is the smallest known livebearing fish species, and one of the smallest overall fish species in the world. It's not often seen for sale these days, although it's quite widely available in hobbyist circles. When buying these, ensure they have been identified correctly as they're sometimes confused with the much more aggressive mosquito fish of the genus Gambusia.