Malaysia

Acanthopsoides spp. are most closely related to the horse-faced loaches of the genus Acantopsis with which they often co-occur in nature, and thus commonly referred to as 'dwarf horse-face loaches'. The genus currently comprises five species of which four were described by Siebert (1991); these were discovered in existing museum holdings...

Acanthopsoides spp. are most closely related to the horse-faced loaches of the genus Acantopsis with which they often co-occur in nature, and thus commonly referred to as 'dwarf horse-face loaches'. Most are very difficult to tell apart but A. robertsi is quite easily identified since it's the only...

Juveniles of this 'species' (probably a group of species - see 'distribution') are traded in large numbers but it's still uncommon to see adult specimens in the hobby suggesting their needs are not catered for often enough. This is a shame as when maintained properly they're hardy, long-lived and interesting aquarium residents. Other trade names include 'long-nosed' and 'banana' loach. Please note although certain that more than one species is being tr...

This is the most commonly-encountered representative of the genus in the aquatic trade though is by no means common. As in other members of the genus body colouration is highly variable, even among individuals collected from a single locality, although a few useful regularities have been observed which can be used to distinguish certain species. This is thought related to the fact that Acrochordonichthys spp. periodically shed their skins and appear paler post-moulting. A. rugosus lends its ...

This species is among the few members of the family to have been exported for the aquarium hobby. It can be differentiated from all other Akysis species by the following combination of characters: head length 17.7-26.5% of standard length (SL); head width 25.4-28.7% SL; body depth at anus 13.0-17.0% SL; caudal peduncle depth 8.1-10.1% SL; interorbital di...

May also be seen on sale under the trade names 'Bala shark' or 'tricolor shark minnow' and perhaps among the most commonly-encountered ornamental fish species. It is unfortunate that the vast majority of those traded are juveniles and tend to come supplied with little to no information provided regarding their potential size and requirements, a fairly ironic situation given the apparently precarious status of wild populations.

An obligate dweller of swiftly-flowing streams and headwaters containing clear, oxygen-saturated water. It often inhabits riffles and runs and is likely to show a preference for shallower zones. Substrates are generally composed of gravel, rocks, boulders or bedrock carpeted with a rich biofilm formed by algae and other micro-organisms. Juveniles are often found in slower-moving stretches with gravel substrate and submerged tree roots. In both cases patches of aquatic plants are only occasionally pres...

This species is not often seen in the aquarium hobby but occasionally exported as bycatch among shipments of other fishes. It was described as a subspecies of the closely-related 'P.' binotatus but is currently considered distinct on a tentative basis as per Ng and Tan (1999) who stated it is 'likely' that the two represent extreme colour forms of a single, variably-patterned species.

This species is so ubiquitous across its natural range that it's often referred to simply as 'common barb'. Basic adult colour pattern consists of a small, dark spot at the base of the dorsal-fin origin and another on the caudal peduncle but overall appearance varies somewhat depending on origin, with the anterior dark spot enlarged ventrally forming a bar or posterior spot extending into the caudal-fin, for example. The spots may also appear darker or lighter in some individuals.

Although sometimes referred to by the vernacular name 'big spot barb' this species is better known in the aquarium hobby as 'clown barb'. It has been widely misidentified as the congener B. everetti, a distinct and uncommonly-traded species from Borneo and thus appears under that name in the majority of available literature. The two are quite easy to tell apart, with B. everetti an overall les...

This species is uncommon in the aquarium hobby, although the scientific name is widely misapplied to Barbodes dunckeri, a similar-looking species native to Peninsular Malaysia and Singapore commonly referred to as 'clown barb' and labelled as B. everetti in the majority of available literature. The two are quite easy to tell apart, with B. everetti an overall less...

This species is rarely-exported for the aquarium hobby but is available on occasion. The name 'P.' kuchingensis is frequently misapplied, however, with subadult forms of 'P.' everetti and some populations of the geographically-variable 'P.' lateristriga regularly traded as such, for example. Though closely-affiliated with and superficially very similar to...

This species is commonly-referred to as 'T-barb' in some countries. It's closely-affiliated with and sometimes traded as the congener B. kuchingensis but that species can be told apart by colour pattern comprising a prominent row of dark spots along the lateral line, plus a short, horizontally-orientated streak extending from the upper part of the operculum. In B. lateristriga the lateral markings usually form a solid stripe and there is no streak extending from the operculum.

The genus Barbonymus was erected by Kottelat in 1999 and contains former members of Barbodes from southeast Asia. The type species is B. schwanenfeldii and currently there exist only three other representatives; B. altus, B. collingwoodii and B. gonionotus. The latter two are rare in the hobby although an SF member has kept B. gonionotus in the past so they are worth looking out for if you harbour an...

This is one of two species traded under the vernacular name 'tinfoil barb', the other being the less well-known congener B. altus. Despite this it appears that B. altus is just as widely available as B. schwanefeldii and in many cases is seen on sale more regularly. Unfortunately both are usually offered at a small size (usually around 50 - 75 mm) with little to no information regarding the eventual size of the fish.

Batasio havmolleri (Smith, 1931) is currently considered a junior synonym of B. fluviatilis. Members of the genus Batasio are characterised by the following combination of characters; laterally-compressed body shape; presence of large sensory pores on the head; a narrow mental region; a pair of prominent posterior proces...

This species is also referred to by the vernacular name 'Malay combtail'. In terms of external characters it differs most obviously from its only congener, B. signata by its overal brownish grey (vs. reddish) body colouration and presence of a mosaic-type patterning in the unpaired fins (vs. absence). The position of relatedness of the genus Belontia with respect to other anabantoids remains somewhat...

Like others in the suborder Anabantoidei this species possesses an accessory breathing organ known as the labyrinth. So-called due to its maze-like structure this organ allows the fish to breathe atmospheric air to a certain extent. Comprising paired suprabranchial organs formed via expansion of the epibranchial (upper) section of the first gill arch and housed in a chamber above the gills, it contains...

Likely to prey on insects and other small invertebrates/zooplankton in nature. Captive fish will normally accept dried products once they're recognised as edible, but should be offered plenty of small live or frozen foods such as Daphnia, Artemia or bloodworm regularly to ensure development of optimal colour and condition. Small insects such as crickets or Drosophila fru...

This species lends it name to the B. bellica group/complex of closely-related species within the genus, an assemblage of which members share the following set of characters: long and slender body with dorsal and ventral margins almost parallel; body depth 23-28 % SL; 30-33 anal-fin rays; 11-13 dorsal-fin rays; 32-34 total vertebrae; body dark brown in colour with iridescent green markings on each individual scale.

B. coccina can be told apart from other members by the following combination of characters: iridescent green lateral blotch present in male, absent in female; body dark red in colour; no parallel vertical iridescent gold opercle bars; pelvic fin falcate with black filamentous tip, remainder red; median fins with discrete iridescent green spots; dorsal fin-rays 9-12; anal-fin rays 27-29; subdorsal scales 7-8; lateral scales 31-32; predorsal scales 18-21. The genus Betta is the most speciose within the family Osphronemidae with almost 70 reco...

Presumed endemic to the geographically-isolated Maliau basin, a roughly circular plateau enclosed by hills in Sabah state, Malaysia, Borneo. The Maliau River is part of the much larger Kinabantangan watershed, Malaysia's second longest river. The type series was collected from heath forest alongside 'Puntius' sealei and Hemibagrus cf. baramensis in acidic water with a pH of 4.2-4.8. Adult individuals were spaced around a metre apa...

The easiest way to distinguish these is by examining the throat markings, which vary according to species, and in the case of B. hipposideros form a black horse-shoe shape. It can also be separated from other group members by the following suite of characters: dark blotch on throat joined to black lower lip; presence of transverse bars in dorsal and caudal fins; absence of dark margin in anal-fin; opercle usually with gold scales, without iridescrent green scales; ventral margin of opercle brown; 6.5-7 subdorsal scales (mode 6.5); 9-10 postdorsal scales (mode 9.5); 31-32 lateral scales (mode 32).

B. ibanorum is uncommon in the hobby and is apparently more frequently fished for food than the aquarium trade. It's included in the Betta akarensis group/complex of closely-related species within the genus, of which members share the following set of characters: possession of pre-orbital and post-orbital stripes (the post-orbital stripe is faint or interrupted in some species); chin bar present; caudal-fin lance...

B. imbellis is included in the Betta splendens complex of closely-related species within the genus, of which members share the following set of characters: head length short 22-31 % SL; often a brightly-coloured body; iris of the eye with iridescent green or blue patches; body elongate or slender; opercles parallel when head viewed dorsally; caudal-fin rays red or brown and contrasting with the iridescent interradial membranes; unpaired fins without an irides...

It can be told apart from other members of the B. pugnax group by the following characters: caudal fin rounded in both males and females; chin bar and second postorbital stripe absent; adult males possess intensive blue or bluish-green iridescent (blackish in preservation) colouration on the sides of the head and throat extending from the snout tip to posterior edge of opercle; possession of two central body stripes, the upper extending from the pec...

It can be told apart from other members of the B. pugnax group by the following characters: chin bar absent; prsence of transverse bars in the dorsal-fin; no transverse bars in caudal-fin; green iridescent scales present on opercle and body; anal fin with iridescent, light blue distal band; a pair black markings below the postorbital stripe on the opercle of females and juveniles; anal-fin rays 25-29 (mode 28); lateral scales 28-30 (mode 29); postdorsal scales 10-11 (mode 12); pelvic-fin length 34.8-50.6% SL; head length 33.4-36.8% SL; inter-orbital width 31.7-37.4% HL.

B. macrostoma is included in the Betta unimaculata complex of closely-related species within the genus, of which members share the following set of characters: body long and slender with depth at dorsal fin origin 18-25 % SL; head large and blunt with width 19-24 % SL; long maxilla and lower lip with distance from tip of lower jaw to posterior end of maxilla 27-54 % HL; caudal-fin rounded in shape, occasionally with elongated median rays; pelvic-fin short and filamentous; dorsal and anal fins relatively pointed.

This species is included in the Betta unimaculata complex of closely-related species within the genus, of which members share the following set of characters: body long and slender with depth at dorsal fin origin 18-25 % SL; head large and blunt with width 19-24 % SL; long maxilla and lower lip with distance from tip of lower jaw to posterior end of maxilla 27-54 % HL; caudal-fin rounded in shape, occasionally with elongated median rays; pelvic-fin short and filamentous; dorsal and anal fins relatively pointed.

B. persephone can be told apart from other members of the group by the following characters: no lateral blotch; body blackish; dorsal surface of head posterior to eye sloping gently; no parallel vertical iridescent gold opercle bars; pelvic-fin falcate with white filamentous tip, remainder black to reddish; median fins covered diffusely with light green iridescence; dorsal fin-rays 9; anal-fin rays 24-27; subdorsal scales 6-7; lateral scales 27-29; predorsal scales 17-18.

B. pi is included in the Betta waseri group of closely-related species within the genus, an assemblage within which members share the following combination of characters: base body colouration clay yellow; opercle scales iridescent gold in mature males of all species except B. tomi; throat with black markings which join with black lower jaw in some species; no chin bar.

It can be told apart from other members of the B. pugnax group by the following characters: chin bar present; no complete second postorbital stripe on opercle; presence of transverse bars in caudal-fin of males; caudal-fin in males broadly lanceolate; opercle scales iridescent blue; anal-fin with reddish subdistal band and black margin; caudal-fin without dark margin; 25-28 anal-fin rays; 28-31 lateral scales; 11-12 postdorsal scales; length of pelvic fins 26.6-43.8 % SL; length of head 27.5-35.2 % SL; inter-orbital width 32.4-53.0 % HL.

Restricted to remnant blackwater habits in heavily-disturbed areas of former peat swamp forest, the original vegetation having been cleared to make space for plantations of oil pal, pineapple and rubber. It's now typically found among overhanging marginal vegetation of irrigation canals, with sympatric species including Belontia hasseltii, Betta bellica, B. imbellis, Sphaerichthys osphromenoides, Parosphromenus sp., and Trichopsis vittata. PH in its natural waters has been recorded over the range 3.9 - 4.2.

At one locality close to Serian, Sarawak, the habitat comprised a swiftly-flowing, clear hill stream with a substrate of quartz gravel and variably-sized rocks. The pH was 7.0 and there was no aquatic vegetation. The stream was approximately 10 metres across at its widest point and between 5-80 cm deep. B. taeniata was found in sluggish stretches among submerged tree roots, leaf litter, or overhanging riparian vegetation.

Within the genus Boraras this species is most similar to B. naevus, though in the latter the red body colouration does not extend into the head. The two also differ in the following characters: anteriormost lateral blotch of comparable size in both sexes in B. maculatus vs. significantly larger in male than female B. naevus; 9-10 + 9 principal caudal-fin rays vs. 9 + 8; 26-29 lateral scales vs. 24-26; i.6.i. pelvic-fin rays vs. i.5.i.

Doubts remain whether B. albolineata is representative of a single species or a group of similarly-patterned, closely-related fish, and the form from Chantaburi province in eastern Thailand was revalidated as B. pulchra (formerly Danio pulcher Smith, 1931) by Kottelat (2013). The status of the currently invalid Danio tweedei (Brittan, 1955) from Kedah state, northern Peninsular Malaysia may also be in question, since fish from that area are clearly...

This species is among the more frequently-traded members of the genus, although confusion regarding its identification and natural distribution is commonplace and it's often misidentified as B. nunus or B. xanthozonus. It can be diagnosed as follows: <30 lateral scales; one black band on head, three on body; first black band on body overlapping first dorsal fin to the extent that almost all fin, except a narrow margin, is black, and beginning opposite centre of opercle...

B. sabanus is very similar to B. doriae and these two are easily confused. However in B. doriae most of the first-dorsal-fin and two-thirds of the pectoral-fin are black while in B. sabanus the last one or two rays of the first dorsal-fin are colourless and less than half of the pectoral-fin is black. In addition there are usually smaller dark 'saddles' betw...

Type locality is '55 miles north of Singapore, freshwater ditches, Mawai District, Johore, Malaysia', and this species has been recorded throughout much of Peninsular Malaysia and southern Thailand. Records from Borneo and Singapore appear to be in error although it may have been extirpated from the latter but as with most members of the genus a degree of confusion surrounds its identity (see 'Notes').

This species is very rare in the trade and is normally imported only as bycatch among shipments of other fishes. It can be told apart from its well-known congener B. dorsiocellata by a combination of characters as follows: lateral line complete (25-30 pored scales vs. 4-9); more scales in the lateral row (29-32 vs. 25-27); larger adult siz...

B. dorsiocellata is sometimes traded as 'green-eyed', 'emerald-eyed' or 'hi-spot' rasbora. There appear to exist several geographical forms of the species at least two of which have been seen in the trade. The most common is the rather plain-coloured fish that is produced commercially in large numbers and has gained popularity due to the bright green/blue reflective patch in the lower part of the eye.

Butis spp. are largely nocturnal ambush predators with cryptic patterning to help them blend in with their surroundings. They can also lighten and darken their body colouration to an extent, have a habit of aligning themselves with solid surfaces whether horizontal, vertical, or inverted, and often swim in an upside-down position. The genus is usually included in the family Eleotridae of which members are often referred to...

This species is relatively uncommon in the aquarium hobby but is available on occasion, sometimes mixed in among shipments of the congener C. salivator. These two can be told apart by the fact that C. salivator possesses a series of distinct dark bars on the head and body in both sexes, a colour pattern unique within the genus. It is also similar t...

It is included in a group often referred to as 'red-eyed puffers', which currently contains four recognised species distributed in Indochina and the Greater Sunda Islands. It can be distinguished from C. borneensis, C. irrubesco, and C. lorteti, the remaining members of this group, by the following combination of characters: 11 dorsal-fin rays; 16 pectoral-fin rays; 11-13 anal-fin rays; 11 caudal-fin rays; eyes not bulging above the head; in males dorsal-fin hyaline, black spot on dorsal-fin base present, pectoral-fin base hyaline, anal-fin hyaline with no black spot at base...

Generally considered to have an enormous natural range extending from Iran to Taiwan and Bali, with records existing from Iran, Iraq, Afghanistan, Pakistan, Nepal, India, Sri Lanka, Bangladesh, Bhutan, China, Myanmar, Thailand, Laos, Cambodia, Vietnam, Malaysia, Indonesia, and Singapore. However there exists significant evidence to suggest that C. gachua as currently understood represents a complex of similar-looking species, and a taxonomic review of the group is clearly required.

Prefers a dimly-lit aquarium with plenty of cover in the form of live plants, driftwood branches, terracotta pipes, plant pots, etc., arranged to form a network of nooks, crannies, and shaded spots. Surface vegetation such as Ceratopteris spp. is also appreciated and makes the fish less inclined to conceal themselves.

This species can be told apart from the similar-looking congener C. baramensis (once considered synonymous with C. melasoma) by absence (vs. presence) of a black spot in the centre of numerous body scales and absence (vs. presence in specimens larger than 120 mm SL) of a barred caudal-fin pattern.

C. oblongus is another name widely misused in the trade but that species has seemingly never been exported and was described as a blueish fish with yellow fins. It's native to streams of Gunung Salak mountain in Bogor Regency, West Java, Indonesia where collecting of ornamental fishes is almost non-existent. Other species of laterally-striped Crossocheilus also exist and may be available from time-to-time but are more easily told apart from the group described above.

In the aquarium trade this species may also be seen on sale as 'skinhead barb'. As with others in the genus little has been written regarding its captive care but it makes a peaceful and unusual addition to larger aquaria. The best way of obtaining it may be to keep an eye on shipments of wild fishes from Indochina and the Greater Sunda Islands as it's rarely imported in large numbers and most often arrives as bycatch.

It can be distinguished from congeners by possessing one or two pairs of barbels, a black blotch at the base of the caudal-fin, and rows of dark spots along the lateral scale rows. Members of Cyclocheilichthys are characterised by possessing a serrated dorsal fin spine, 9 branched pelvic fin rays, a conical snout, small, subterminal...

This species is currently understood to have a considerable natural range extending eastwards from Myanmar via Thailand, Laos, Cambodia as far as Vietnam and to the south through Peninsular Malaysia and into the Greater Sunda Islands of Borneo and Sumatra. It thus occurs in the Mekong, Mae Klong and Chao Phraya drainages as well as a host of other river basins, lakes and reservoirs including the Tonlé Sap river...

An ideal resident of the larger, well-furnished community set-up though it might upset slow-moving or timid companions simply by its size. There are plenty of suitable tankmates including many cyprinids, loaches, cichlids, catfish and characins, but as always when selecting a compatible community of fish proper research is essential. A community based around one of its native countries or river basins...

D. polota is sometimes referred to as D. quadrifasciatus (Sevastianov, 1809) but the original name of the latter, Chaetodon quadrifasciatus Sevastianov, 1809, is a junior primary homonym of the older Chaetodon quadrifasciatus Bloch & Schneider 1801, thus Coius polota Hamilton, 1822 takes precedence.

Native to parts of southern Borneo, eastern Sumatra, Singapore, and the Malay Peninsula. In Singapore it's considered highly-endangered and is now restricted to the Central Catchment nature reserve. Type locality is 'Tuluk and Gunung Sahilan, Sumatra, Indonesia' which appears to correspond to localities within the Kampar River drainage of which the former is now known as Teluk Meranti and is o...

D. johorensis can be distinguished by the following combination of characters: 5-6 dark, lateral stripes on body in specimens larger than 40 mm; stripes immediately above and below the central stripe (stripes '+1' and '-1', respectively) located on scale rows +2 and -2, not touching dorsal and ventral midlines (except in some small specimens from Peninsular Malaysia); stripes broad, typically between 0.5-1 scale rows deep...

This species is rare in the aquarium trade with the majority of fish available under the name or the vernacular 'pentazona barb' being the closely-related D. hexazona. The two are almost identical in terms of both appearance and meristics but can be separated by the fact that D. pentazona possesses a small dark marking at the posterior base of the dorsal-fin which is absent in D. hexazona.

This species is one of three similar-looking members of the genus alongside D. johorensis and D. trifasciatus, while it may also be confused with Striuntius lineatus. Juveniles of all except S. lineatus exhibit a vertically-barred rather than laterally-striped colour pattern, the metamorphosis to adult patterning beginning at around 20 mm SL and normally being complete by 30-40 mm.

Probably restricted to western and southern Thailand and northern Peninsular Malaysia plus some small surrounding islands. In Thailand records exist from Surat Thani (Ko Samui island), Nakhon Si Thammarat, Ranong, Trang, Phuket, Ratchaburi, Kanchanaburi, and Mae Hong Son provinces while in Peninsular Malaysia it's been collected from Perak and Kedah states, including th...

Appears to have a disjunct range with records from Pahang state in central Peninsular Malaysia, plus the Mae Klong and Chao Phraya river systems in Thailand, but it has not been recorded in Peninsular Thailand. Type locality is 'Pahang River, Kuala Tembeling, eastern slope of Sangka Dua Pass, Malaysia, elevation about 2000 feet'.

E. octozona can be told apart from congeners by the following combination of characters: bar 1 present; presence of a large and distinct black mark anterior to anus, visible in lateral and ventral views; simple dorsal-fin rays black, sometimes extending onto first branched ray; other rays hyaline or with diffused chromatophores on last dorsal-fin ray, adjacent to upper extremity of bar 6; 25-31 serrae on last simple dorsal-fin ray; uppermost ray of pectoral-fin with faint black margin; width of bar 5 equal to 1-1½ lat...

Confusion with similar-looking fishes from other genera, such as Garra cambodgiensis, laterally-striped Crossocheilus spp. or Gyrinocheilus aymonieri is not uncommon, and are largely attributable to the use of trade names such as 'false flying fox' or 'Siamese flying fox'. When compared with the other species E. kalopterus exhibits several unique characters but perhaps the simplest way to identify it is by the characteristic white-edged, red and black coloured fins and the presence of two pairs of barbels.

There currently exist a dozen described Esomus spp. alhough few are seen in the aquarium trade and none are especially popular. Most are commonplace in their native countries and also quite plainly-patterned so generally overlooked by collectors. Their most common use in some areas is actually as a feeder fish in the aquaculture of larger species. They're characterised by greatly-enlarged pectoral fins and two pairs of barbels, of which the maxillary pair are extremely long and usually reach the pectoral fins.

Also known by the vernacular names ''black band garra', 'stone-lapping minnow' or 'false flying fox' and continues to be traded as Garra taeniata, an invalid name synonymous with G. cambodgiensis. It is often confused with those laterally-striped members of the genus Crossocheilus sold as 'Siamese algae eater' or 'Siamese flying fox', but in those species the dark lateral body stripe continues into the caudal-fin whereas in G. cambodgiensis it terminates...

Field observations have revealed that individuals almost always position themselves facing into the flow, either along the sides, behind or under rocks, their specialised morphology (see 'Notes') allowing them to feed and maintain position without being swept away. Not many fishes have been able to successfully colonise this narrow ecological niche, but syntopic species in nature include Gastromyzon extrorsus (Petagas River), G. introrsus, G. lepidogaster (Pad...

G. cornusaccus can be distinguished from congeners by the following combination of characters: secondary rostrum present; incomplete postoral pouch present reduced to pockets located posterior to mouth corners; sublacrymal groove present and visible when viewed laterally; juveniles with thick, cream-coloured bars and blotches on body, adults uniformly-coloured; head dorsum plain; subopercular groove absent; gill slit vertical; snout truncate when viewed dorsally; no scales on abdomen; 55-61 lateral line scale...

One of the more commonly-traded members of the genus and often found in mixed shipments which may contain other Gastromyzon spp. or related fishes like Beaufortia kweichowensis, these often labelled collectively as 'Borneo sucker', 'Hong Kong pleco', 'butterfly loach', etc. It's sometimes misidentified as G. punctulatus, a species not currently traded which possesses yellow finnage and a lighter-coloured, less-intensely spotted body.

This species has probably not been seen in the aquarium trade yet but has been maintained by a few private collectors. It can be distinguished from congeners by the following combination of characters: secondary rostrum present; complete postoral pouch present; body colour brown with up to 9 thin gold stripes; head dorsum dark brown with gold reticulate pattern; subopercular groove absent; gill slit vertical; sublacrymal groove present; sno...

In fact G. farragus and G. ocellatus represent one of 12 pairs of cryptospecies to be found in the genus, differing in subtle aspects of patterning. Cryptospecies are defined as morphologically similar, but reproductively isolated species which in fishes often inhabit adjacent river basins but in some cases occur sympatrically. The phenomenon may be a result of parallel evolution, and is not normally considered to represent an early stage of speciation.

The type locality was a fast-flowing, shallow (50 - 80 cm deep) river between 5-10 metres in width with a substrate of gravel and large rocks. PH was measured at 7.7 and the habitat was unshaded with on...

This species is variable in terms of colour. Large adults may be bright green, orange or barred but are apparently capable of changing from barred to plain in just a few seconds. These different forms have been recorded in the same natural habitats, and some individuals also possess prominent nuptial tubercules covering much of the body.

Field observations have revealed that individuals almost always position themselves facing into the flow, either along the sides, behind or under rocks, their specialised morphology (see 'notes') allowing them to feed and maintain position without being swept away. In nature G. monticola occurs sympatrically alongside Garra borneensis, Osteochilus ingeri, Gastromyzon auronigrus, G. cornusaccus, Nemacheilus olivaceus,...

Some individuals do appear very similar to G. farragus, with noticeable spotting on the head, but can usually still be told apart by examining the caudal-fin which tends to contain only a single thick, dark vertical bar in G. farragus whereas in G. ocellatus there are more often two bars, one thick, one thinner. Intermediate forms do exist though meaning identification is sometimes tricky.

It can be distinguished from congeners by the following combination of characters: gill slit angular; presence of subopercular groove, running continuous to origin of pectoral-fin; body black with numerous, small, evenly-spaced light brown spots; head dorsum black with numerous cream spots; pectoral and pelvic fins with cream spots; caudal-fin with iridescen...

One of the more commonly-traded members of the genus and often found in mixed shipments which may contain other Gastromyzon spp. or related fishes like Beaufortia kweichowensis. These are typically labelled 'Borneo sucker', 'Hong Kong pleco', 'butterfly loach', etc. regardless of species. It's sometimes misidentified as G. punctulatus, a species not currently traded which possesses yellow finnage and a lighter-coloured, less-intensely spo...

Many loaches from the families Nemacheilidae, Balitoridae and Gastromyzontidae are also suitable but research your choices before purchase in order to be sure as some are excessively aggressive or otherwise competitive. Gastromyzon spp. tend to exist in lo...

G. filamentosa has appeared in the ornamental trade but is evidently delicate and therefore recommended only to experienced aquarists. It can be told apart from congeners by the following combination of characters: possession of filamentous dorsal and anal-fin rays in males; body transparent to translucent yellowish-brown colour in life; absence of a highly-extended second anal-fin spine; absence of carmine-red pigmentation in the dorsal, anal an...

Gymnothorax tile

The pinkish-white form of this species is among the most widely-traded aquarium fishes in the world but does not occur naturally and is in fact a leucistic variant bred specifically for ornamental purposes. The 'short-bodied' or 'balloon' variety also continues to be fashionable but under no circumstances do we recommend the purchase of these intentionally disfigured animals which tend to have much shorter life spans than the naturally-shaped fish. There also exists a less-po...

The reason that the paired fins are orientated vertically, as opposed to horizontally in other members of the genus, is unclear but Roberts (1989) suggest it may be an adaptation to this species' ecology, in that it displays a preference for submerged vegetation or woody structures as opposed to rocky environments. They're certainly adapted for a rea...

This species appears is rare in the aquarium hobby with the majority of images both online and in the published literature instead depicting H. confuzona or H. parclitella. All three are members of the H. ocellata 'group' within the genus which currently contains H. ocellata, H. bilineata, H. confuzona, H. orthogoniata, H. ogilviei and H. parclitella. Following Tan and Ng (2005) these share possession of...

The water at the type locality is described as 'clear and fast-flowing over a rocky substratum'. H. parclitella was observed in shallow water among the rocks, and sympatric species included Garra cambodgiensis, Neolissochilus soroides, Poropuntius smedleyi, 'Puntius' binotatus, Homaloptera nebulosa and Amblyceps mangois.

The genus Homalopteroides was revalidated by Randall and Page (2012) and is told apart from the related Homaloptera by the following character combination: dorsal-fin origin above pelvic-fin; ≤ 60 lateral-line scales; ≤ 30 predorsal scales; oral morphology consisting of two thin and widely separated rostral barbels on each side of the mouth, thin crescent-shaped lips, the absence of any structure such as a mental pad or lobes between the lateral portions of the lower lip, and a chin that extends anterior to the lateral portions of the lower lip.

Described from close to the village of Ban Khiri Wong, Nakhon Si Thammarat Province, southern (peninsular) Thailand but since recorded throughout much of the country, including the Chao Phraya and Mekong river systems. It's also known from Peninsular Malaysia and parts of the Mekong basin in Laos, Cambodia and Vietnam. Rainboth (1996) considered it 'probably the commonest member of the genus in the Mekong' and in a 2008 study of balitorid habitats in centra...

Tan (2009) redescribed H. stephensoni and it possesses the following unique character set: sub-inferior mouth with thin barbels; five principle, 12-13 secondary pectoral fin rays (usually 12); scaleless belly; pattern of keeled scales on anterior, dorsal portion of body consisting of a single central keel in the posterior third of each scale; 43-50 lateral line...

Described from Kota Tinggi district, Johor state, Peninsular Malaysia but since recorded throughout much of the country as well as parts of the the Mekong river system in Thailand, Laos, Cambodia and Vietnam plus the Kapuas and several other river basins in Kalimantan Barat (West Kalimantan) province, Indonesian Borneo. In the Kahang River, part of the Endau drainage in Jo...

An obligate dweller of swiftly-flowing streams and headwaters containing clear, oxygen-saturated water. It often inhabits riffles and runs and is likely to display a preference for shallower zon...

It is very similar to H. wetmorei but can be told apart by possessing yellow (vs. orange to red in H. wetmorei) pectoral, pelvic, and anal fins. It thus appears likely that yellow-finned fish marketed as H. wetmorei or 'lemon fin barb' in the ornamental trade are actually H. vernayi, with both species available on a regular basis.

H. wetmorei is distinguished from congeners by the following combination of characters: upper body reddish brown; robust body, not compressed; usually 4½ scale rows between lateral line and dorsal-fin origin; 8-12 rakers on first gill arch; 7-9 predorsal scales; 14 circumpeduncular scale rows; usually two scale rows separating vent and anal-fin; <29 lateral line scales; 9-14 serrations on the spinous dorsal-fin ray; distance between distal serrae on posterior margin of last unbranched dorsal-fin ray much greater than the width of their bases.

The type series was collected from a blackwater stream in Toh Daeng Peat swamp forest with the fish found in slow-moving or standing habitats among submerged shoreline roots and leaf litter. In the Tapi River floodplain it was collected from stagnant waters, one of which was an oxbow lake approximately 200 m long and 30 m across at its widest point, surrounded by 'freshwater scrub and swamp trees'. The substrate was composed of mud, woody structures including fallen twigs and branches, and decomposing plant material.

The genus Kottelatlimia was erected by Nalbant (1994) to accommodate K. katik which had previously been assigned to Lepidocephalichthys but differs in various aspects noted earlier by Kottelat and Lim (1992) including: relatively small adult size; scaleless...

Generally peaceful though it may predate on smaller fishes and is somewhat timid so does not compete well with much larger, robust or otherwise boisterous species. Peaceful, comparably-sized cyprinids, loaches and other catfishes perhaps constitute the best options but be sure to research your choices thoroughly prior to purchase.

Known from the Malay Peninsula and Singapore plus Greater Sunda Islands of Sumatra, Borneo and Java in Indonesia with populati0ns from Thailand, Cambodia, Laos and Vietnam formerly considered as conspecific now referred to K. geminus (Ng, 2003). Type locality is given as 'Bandjarmassing' which corresponds to a town now more commonly referred to as 'Banjarmasin' in South Kalimantan (Kalimantan Selatan) province, Indonesia (Borneo).

Type locality is 'Palembang, Sumatra, Indonesia', but this species is widely-distributed throughout much of Southeast Asia including major river systems in Thailand, Laos, Cambodia, Vietnam and Peninsular Malaysia plus the Greater Sunda Islands of Sumatra, Borneo and Java. It's been extensively recorded from the Mekong, Cha...

Has been recorded in fast-flowing environments but most commonly associated with peat swamp forests and associated blackwater streams. Many such habitats have suffered degradation of some kind but in unaltered cases the dense canopy of branches above means very little light penetrates the surface of such environments, and riparian vegetation also tends to grow thickly.

Different populations vary in appearance somewhat (see image of Salween specimen for example), and L. leptocheilus may turn out to represent a group of closely-related species rather than a single taxon. The population from the Cambodian Mekong has been considered to represent a distinct species, Labiobarbus lineatus, although that name is currently a synonym of L. leptocheilus following Kottelat (2013). It is widely used in the ornamental trade, however.

It can be distinguished from congeners by the following combination of characters: 61-68 scales in the lateral series; a small, well-defined, sometimes ocellated black spot on the shoulder and another in the centre of the caudal peduncle; body without longitudinal stripes formed by spots on scales; caudal fin uniformly dusky or colourless, lobes without stripes or black margins.

Considered synonymous with the congener L. laubuca for a number of years, thus reports of that species from anywhere in Indochina actually refer to the current concept of L. siamensis. Given the distribution of L. siamensis, it seems likely that many of the fish entering the aquarium trade are also this species and not L. laubuca. The two species can be distinguished by...

This species is not traded often but is occasionally available as bycatch among shipments of other species. It can be told apart from congeners most easily by the prominent, dorsally-orientated, roughly semi-circular lamina circularis in males, a feature unique within the genus. Other distinguishing characters include a combination of: forked caudal-fin; relatively small adult size; dark <-shaped mark at the base of the caudal-fin; a black spot at the base of branched caudal-fin rays 3-5.

At one locality close to Ban Na Hwai, Chiang Mai Province, Thailand L. hasselti was collected from a shallow (<0.5 m) pool in a swampy zone between forest and rice fields. The substrate was of mud, there was no aquatic vegetation and it was being used as drinking water by local animals, with the only effluent connected to a 'small creek'. Other species found there were Physoschistura pseudobrunneana, Rasbora hobelmani and Systomus cf. orphoides.

This species is not traded often but may be available occasionally bycatch among shipments of other species. It can be told apart from congeners most easily by the following combination of characters: truncate caudal-fin; scaleless patch on top of head; caudal-fin with reticulated patterning, sometimes appearing as a series of thin vertical bars; body reddish brown with darker markings; 3-7 predorsal and 3-6 postdorsal thin, transverse dark bars running across the dorsal surface; relatively deep...

The distribution records for this species are somewhat confusing. It was described from the confluence of the Lamatang and Enim rivers, Palambang Province, Sumatra and has since been recorded from Thailand (Chao Phraya River), Peninsular Malaysia (Pahang River) plus the islands of Java (Solo River) and Borneo (Kapuas and Barito rivers).

Lobocheilos species are specialised grazers of periphyton, benthic algae, and other organisms which grow attached to rocks and other solid surfaces, and they typically leave visible scrape marks in places where they have been feeding. They are by no means herbivorous and should be offered meaty foods such as live or frozen chironomid larvae (bloodworm), Tubifex, Artemia, chopped prawn, etc., along with good quality, sinking dried products, at lea...

There is a fish of unknown geographical origin which matches the majority of diagnostic features for L. setigerum with the exception that the dark lateral stripe is absent in the anterior portion of the body and is not composed of interconnected spots. It is relatively common in the aquarium trade and included here as L. cf. setigerum until a confirmed identity is established.

This species is occasionally marketed as 'Burmese Rainbow Barb' in the ornamental trade. others may have been exported in small numbers or as bycatch among shipments of other species. It was referred to as Mystacoleucus marginatus for decades, but that name is a simultaneous subjective synonym of M. obtusirostris following Kottelat (2013).

This species was originally named Nemacheilus masyae but following ICZN rules the spelling was later corrected to N. masyai because it's named after a man. It can be distinguished from the majority of congeners by body patterning comprising 14-18 short, dark vertical bars on each flank, 12-17 saddle-like markings running along the dorsal surface, a dark spot on the caudal peduncle at the termination of the lateral line and a dark blotch in the lower half of the first few dorsal-fin rays.

Images depict clear, tea-coloured forest streams with dense marginal vegetation and substrates of sand and/or small rocks and pebbles. It's also known from environments with sandy substrate and organic debris in the form of submerged roots/branches and le...

N. selangoricus is most similar to N. spiniferus since it possesses acuminate scales above and below the lateral line on the caudal peduncle, a feature not shared with any other congener. It can however be distinguished by the following characters: presence of 8-12 very regular dark bars on flanks (vs. 10-13 irregularly-shaped bars in N. spiniferus), 3-5 times...

It's a member of the N. selangoricus group of species within the genus, an assemblage first recognised by Hadiaty and Kottelat (2009) and characterised by possession of two rows of horizontally-arran...

We're unsure if this species has ever been traded on a commercial basis though we know of several private collectors who've maintained it. It's difficult to confuse with any other balitorid due to the small adult size and distinctive orange to reddish-brown body colouration, and was originally described as a subgenus by Herre (1944) before Silas (1953) raised it to full genus status. It was later syonymised with Homaloptera by Alfred (1969) but has been generally accepted as a distinct, valid genus since Roberts (1989).

This species is widely-distributed throughout Peninsular Thailand, Malaysia (Malay Peninsula plus the states of Sabah and Sarawak, Borneo), Singapore and Indonesia, with records from the Riau Archipelago, Sumatra, Java, Borneo, Bali, Lombok, and Sulawesi existing in the latter. Type locality is 'Panimbang River, Perdana, Indonesia'. It's typically found in coastal, normally brac...

One of a handful of Pangio spp. traded under the generic name 'eel loach'. Wild populations tend to differ in colour pattern to some extent, with some exhibiting a more intense lateral stripe while others display an irregular pattern of fine dark spots on the body, for example.

Infrequently encountered in the aquarium trade, most often as bycatch among shipments of similar-looking congeners. It's included in the P. kuhlii group of closely-related species within the genus, and can be distinguished from other members by...

One of a handful of Pangio spp. traded under the generic name 'eel loach', though it's most commonly available as bycatch among shipments of other fishes. Some authors have suggested it may be synonymous with P. anguillaris bu...

It's included in the P. kuhlii-oblonga group of closely-related species within the genus as per Bohlen et al. (2011) and can be...

Originally known only from Johor and Pahang states, Peninsular Malaysia but since recorded from the Batang Hari river drainage, Sumatra and the Kapuas system, Kalimantan Barat (West Kalimantan) province, Indonesian Borneo. Type locality is 'Tahan River, Pahang, Malaysia'.

Described from close to the city of Bogor, Jawa Barat (West Java) province on the island of Java, Indonesia, but currently considered widespread with recorded occurrences on Java, Sumatra, Borneo and mainland Peninsular Malaysia. Some populations exhibit lighter...

Some Sumatran and Bornean populations of the highly variable P. shelfordii superficially resemble P. piperata which can cause issues with identification as the two often occur together. In general however the body patterning in P. piperata is than that of P. shelfordii and they differ in vertebral counts (mode 47 vs. 50, respectively).

This species is among the most frequently-misidentified in the hobby, almost always being traded as P. kuhlii. However the latter is currently considered endemic to Java, from where commercial collections are almost unheard of, and has possibly never been seen in aquaria.

Patterning is highly variable depending both within and between populations and the species may comprise an assemblage containing a number of taxa (Kottelat and Lim, 1993; Tan and Kottelat, 2009). For example, those from Singapore and Johor possessing intricate mottling on the dorsal surface while individuals from Terengganu have a series of saddle-like markings either alternating or connecting with the midlateral markings.

According to Kottelat (1990) the chief distinguishing characters for the genus are: possession of 9-18½ branched dorsal-fin rays; colour pattern without black basal caudal bar, but with an ocellus at upper base of caudal-fin; upper lip with or without several rows of papillae; lower lip with a broad median interruption and a widened, papillated pad on each side.

Type locality is given only as 'Sarawak state, Borneo, East Malaysia', with the material deriving from Sibu Division in western Sarawak. It's since been recorded at other localities within Sarawak, to which it appears endemic, but it remains unclear whether all represent a single species or not with those from the area around Lundu in northwestern Kuching Division in par...

Parosphromenus spp. have been grouped arbitrarily based on courtship behaviour in males which adopt a 'head-down', 'head-up', or 'horizontal' position depending on species. P. harveyi belongs to the former, most abundant, assemblage in which the male assumes a near-vertical position with the head lowermost and fins splayed during nuptial displays.

In males the dorsal, anal and caudal have a black subdistal band bordered on either side by bluish iridescent bands, with the inner one significantly thicker. The pelvic fins are iridescent emerald green with a dark blotch at the base and a subdistal dark spot, while the pectoral fins are hyaline.

This species has been known in the aquarium hobby since the mid-twentieth century but has almost always been identified as the congener P. deissneri and appears as such in the majority of aquarium literature. It can be told apart from congeners by the colour pattern in adult males (see 'Sexual Dimorphism') plus possession of 11-12 spinous dorsal-fin rays, 13-14 spinous anal-fin rays and 6-8 segmented anal-fin rays.

Type locality is given as 'Kuala Tahan, Pahang (King George V National Park)' which corresponds to the village of Kuala Tahan in Pahang state, central Peninsular Malaysia. The settlement is located at the confluence of the Tahan and Tembiling Rivers which form part of the Pahang river basin, while the national park was re...

This species occasionally appears in the aquarium hobby, but the trade is largely reliant on commercially-produced 'tiger barbs' of questionable origin. It is distinguished from congeners by the following combination of characters: lateral line complete; 14 circumpeduncular scales; 21-23+2 scales in the lateral row; dorsal-fin black with red outer band; pelvic fins red.

Recorded from numerous river basins in Vietnam, Cambodia, Laos, Thailand, and Peninsular Malaysia, including the Mekong, Chao Phraya, Mae Khlong, Chanthaburi, Tapi, Golok, Terengganu, Pahang, Endau, and Muar. Type locality is 'Kratt, southeastern Siam', which corresponds to modern-day Trat Province, Thailand.

One of four similar-looking, laterally-striped 'Puntius' spp. native to Southeastern Asia alongside 'P.' gemellus, 'P.' johorensis, and 'P.' trifasciatus. Of these 'P.' lineatus is most often confused with 'P.' johorensis by virtue of the fact that these two seem most common in the aquarium trade. Telling them apart is relatively simple since 'P.' lineatus has 0 - 1 pairs of barbels (vs. 2 pairs in the other three species), juv...

The identity of this species was unclear for a number of decades prior to its redescription by Ng and Kottelat (2013). The confusion originated with Brittan (1954) who misidentified specimens of R. tornieri as R. dusonensis and was exacerbated by Alfred (1963) who concluded that the holotype of R. dusonensis was conspecific with R. myersi.

Type locality is 'Central Sumatra, Indonesia', with additional records existing from Cambodia (Mekong drainage), Peninsular Malaysia (Perak River), Sumatra (from the Siak River, Riau province to the Musi River, South Sumatra (Sumatera Selatan) province) and Borneo (south and westwards from the Belait river basin in Brunei Darussalam to the Sambas drainage in West Kalimantan (Kalimantan Barat) province, Indonesia and probably in Sarawak, Malaysia).

Should not prove difficult to maintain under the correct conditions. We strongly recommend keeping it in a tank designed to resemble a flowing stream or river with a substrate of variably-sized rocks, sand, fine gravel, and some water-worn boulders. This can be further furnished wit...

The genus Sphaerichthys currently comprises four species with S. osphromenoides by far the best known in the hobby. It's easily distinguishable from the congeners S. vaillanti and S. acrostoma since both exhibit a notably more elongate head and body profile, reverse sexual dimorphism (i.e. females are the more colourful/strongly-patterened), and are paternal mouthbrooders.

This species is available occasionally, though not always under the correct name. We've seen it on sale as S. elegans on numerous occasions, for example, and other trade names include 'Sumatran blue neon goby' (it's not found on Sumatra), 'cobalt blue goby' and 'freshwater neon goby'. Further...

The genus Puntius is currently viewed as something of a catch-all for well over 100 species of small to mid-sized cyprinid. Most experts agree that a full revision is required with the likely outcome that many species will be placed into new or different genera since it is clearly polyphyletic. When describing the grouping in 1822 Hamilton identified the defining characteristics as: presence or absence o...

The common name is derived from the ability of genus members to produce audible sounds via a specialised pectoral mechanism which is unique within the family Osphronemidae. The structure comprises modified pectoral-fin tendons and muscles which are stretched and plucked by basal elevations of two fin rays in a similar way to guitar strings via rapid beating of the fins. The pectoral-fins beat alternately, each able to generate short or long bursts of sound. These sounds are produced by both sexes, predom...